@import"https://fonts.googleapis.com/css2?family=Fraunces:ital,opsz,wght@0,9..144,400;0,9..144,600;0,9..144,900;1,9..144,400&family=Spline+Sans:wght@400;500;600&display=swap";
:root{--ink:#1a1f1a;--ink-soft:#3d453c;--paper:#f4f0e6;--paper-warm:#ece5d3;--card:#fbf9f2;--line:#d4cdb8;--moss:#2f4a32;--moss-bright:#4a7a4f;--rust:#b5562a;--gold:#c8923a;
--sf-cerambycinae:#8B4513;--sf-lamiinae:#2F5233;--sf-lepturinae:#1E3A5F;--sf-prioninae:#4A2040;--sf-spondylidinae:#3D3D20;
--serif:"Fraunces",Georgia,serif;--sans:"Spline Sans",system-ui,sans-serif;--maxw:1180px;--shadow-lift:0 8px 28px rgba(26,31,26,.16)}
*{box-sizing:border-box;margin:0;padding:0}html{scroll-behavior:smooth}
body{font-family:var(--sans);color:var(--ink);background:var(--paper);line-height:1.6;-webkit-font-smoothing:antialiased;background-image:radial-gradient(circle at 12% 8%,rgba(74,122,79,.05),transparent 40%),radial-gradient(circle at 88% 92%,rgba(139,69,19,.04),transparent 40%)}
a{color:var(--moss);text-decoration:none}a:hover{color:var(--rust)}img{max-width:100%;display:block}
.wrap{max-width:var(--maxw);margin:0 auto;padding:0 1.5rem}
.site-header{border-bottom:1px solid var(--line);background:#f4f0e6f0;backdrop-filter:blur(8px);position:sticky;top:0;z-index:50}
.site-header .wrap{display:flex;align-items:center;gap:2rem;padding:.85rem 1.5rem}
.brand{font-family:var(--serif);font-weight:900;font-size:1.15rem;letter-spacing:-.01em;color:var(--ink);white-space:nowrap}
.brand:hover{color:var(--moss)}.brand .mark{color:var(--rust)}
.nav{display:flex;gap:1.35rem;flex-wrap:wrap;font-size:.9rem}
.nav a{color:var(--ink-soft);font-weight:500}.nav a:hover{color:var(--rust)}
.nav .pill{background:var(--moss);color:var(--paper);padding:.3rem .7rem;border-radius:2px}.nav .pill:hover{background:var(--rust);color:var(--paper)}
.hero{padding:4.5rem 0 3.5rem;background:var(--moss);color:var(--paper)}
.hero .eyebrow{color:var(--gold)}
.hero h1{font-family:var(--serif);font-weight:900;font-size:clamp(2.6rem,6vw,4.6rem);line-height:1.02;letter-spacing:-.025em;max-width:16ch;color:var(--paper);margin-top:.4rem}
.hero h1 em{font-style:italic;color:#a8c49a}
.hero .lede{margin-top:1.1rem;font-size:1.15rem;color:#cdd6c8;max-width:52ch}
.hero-cta{margin-top:1.8rem;display:flex;gap:.8rem;flex-wrap:wrap}
.hero .btn-primary{background:var(--gold);border-color:var(--gold);color:#2c1d08}.hero .btn-primary:hover{background:#dba74a}
.btn{display:inline-block;font-weight:600;font-size:.92rem;padding:.7rem 1.3rem;border-radius:2px;border:1.5px solid var(--moss);transition:transform .12s,background .15s}
.btn-primary{background:var(--moss);color:var(--paper)}.btn-primary:hover{background:var(--ink);color:var(--paper);transform:translateY(-1px)}
.btn-ghost{background:transparent;color:var(--moss)}.btn-ghost:hover{background:var(--moss);color:var(--paper)}
.stats{display:grid;grid-template-columns:repeat(auto-fit,minmax(110px,1fr));gap:1px;background:var(--line);border:1px solid var(--line);margin:2.5rem 0}
.stats .stat{background:var(--card);padding:1.3rem 1rem;text-align:center}
.stats .num{font-family:var(--serif);font-weight:900;font-size:1.9rem;line-height:1;color:var(--moss)}
.stats .lbl{font-size:.72rem;text-transform:uppercase;letter-spacing:.09em;color:var(--ink-soft);margin-top:.35rem}
.section{padding:3rem 0}
.eyebrow{font-size:.73rem;text-transform:uppercase;letter-spacing:.14em;color:var(--rust);font-weight:600}
.section h2{font-family:var(--serif);font-weight:900;font-size:clamp(1.7rem,3.5vw,2.5rem);letter-spacing:-.02em;margin:.3rem 0 1.4rem}
.grid{display:grid;grid-template-columns:repeat(auto-fill,minmax(230px,1fr));gap:1rem}
.card{background:var(--card);border:1px solid var(--line);border-radius:3px;overflow:hidden;transition:transform .14s,box-shadow .14s;display:flex;flex-direction:column}
.card:hover{transform:translateY(-3px);box-shadow:var(--shadow-lift)}
.subfam-strip{height:5px;width:100%;display:block}
.card-body{padding:.85rem .9rem 1rem;flex:1;display:flex;flex-direction:column}
.card .sci{font-family:var(--serif);font-style:italic;font-weight:600;font-size:1.02rem;line-height:1.2}
.card .common{font-size:.82rem;color:var(--ink-soft);margin-top:.2rem}
.card .meta{margin-top:auto;padding-top:.6rem;font-size:.72rem;color:var(--ink-soft);display:flex;gap:.5rem;flex-wrap:wrap;align-items:center}
.tag{display:inline-block;font-size:.68rem;font-weight:600;text-transform:uppercase;letter-spacing:.05em;padding:.18rem .5rem;border-radius:2px;background:var(--paper-warm);color:var(--ink-soft)}
.tag.iucn{background:var(--moss);color:#fff}.tag.iucn.threatened{background:var(--rust)}
.tag.pollinator{background:#2a6b3a;color:#fff}.tag.eppo{background:#8B0000;color:#fff}
.crumb{font-size:.8rem;color:var(--ink-soft);padding:1.1rem 0;border-bottom:1px solid var(--line)}
.crumb a{color:var(--moss)}
.genus-head{padding:2.5rem 0}
.genus-title{font-family:var(--serif);font-weight:900;font-size:clamp(2rem,4.5vw,3.2rem);font-style:italic;line-height:1.05;letter-spacing:-.02em}
.genus-common{font-size:1.2rem;color:var(--moss);font-weight:600;margin-top:.3rem}
.genus-authorship{color:var(--ink-soft);font-size:.95rem;margin-top:.2rem}
.badge-row{display:flex;gap:.5rem;flex-wrap:wrap;margin-top:1rem}
.factgrid{display:grid;grid-template-columns:repeat(auto-fit,minmax(150px,1fr));gap:1px;background:var(--line);border:1px solid var(--line);margin:1.5rem 0}
.factgrid:empty{display:none}
.fact{background:var(--card);padding:.9rem 1rem}
.fact .k{font-size:.68rem;text-transform:uppercase;letter-spacing:.08em;color:var(--ink-soft)}
.fact .v{font-weight:600;font-size:.98rem;margin-top:.2rem}
.panel{background:var(--card);border:1px solid var(--line);border-radius:4px;padding:1.5rem;margin:1.3rem 0}
.panel h3{font-family:var(--serif);font-weight:900;font-size:1.3rem;margin-bottom:.7rem}
.panel p+p{margin-top:.7rem}
.pheno{display:grid;grid-template-columns:repeat(12,1fr);gap:3px;margin-top:.8rem}
.pheno .m{text-align:center;padding:.5rem .1rem;font-size:.68rem;font-weight:600;background:var(--paper-warm);color:var(--ink-soft);border-radius:2px}
.pheno .m.active{background:var(--moss);color:var(--paper)}.pheno .m.active.peak{background:var(--rust);color:var(--paper)}
.host-list{display:flex;flex-wrap:wrap;gap:.4rem;margin-top:.7rem}
.host-item{font-family:var(--serif);font-style:italic;font-size:.88rem;background:var(--paper-warm);border:1px solid var(--line);padding:.2rem .7rem;border-radius:2px}
.cons-panel{border-left:4px solid var(--rust);background:#b5562a0d;padding:1rem 1.2rem;border-radius:0 4px 4px 0;margin:1rem 0}
.taxon-list{list-style:none}
.taxon-list li{border-bottom:1px solid var(--line);padding:.7rem 0;display:flex;justify-content:space-between;gap:1rem;align-items:baseline}
.taxon-list .sci{font-family:var(--serif);font-style:italic;font-weight:600}
.taxon-list .aux{font-size:.8rem;color:var(--ink-soft)}
.sfam-tile{display:block;border-radius:4px;padding:1.1rem 1rem;transition:transform .12s ease,box-shadow .12s ease;color:#fff;text-decoration:none}
.sfam-tile:hover{transform:translateY(-3px);box-shadow:var(--shadow-lift);color:#fff}
.sfam-tile .sf-name{font-family:var(--serif);font-weight:900;font-size:1.1rem;line-height:1.1}
.sfam-tile .sf-count{font-size:.82rem;margin-top:.3rem;opacity:.85}
.sfam-grid{display:grid;grid-template-columns:repeat(auto-fit,minmax(160px,1fr));gap:.7rem;margin-top:1.4rem}
.search-box{width:100%;font-size:1.05rem;font-family:var(--sans);padding:.9rem 1.1rem;border:1.5px solid var(--line);border-radius:3px;background:var(--card)}
.search-box:focus{outline:2px solid var(--moss-bright);border-color:var(--moss)}
.filter-chip{font-size:.8rem;font-weight:500;padding:.4rem .85rem;border-radius:999px;border:1.5px solid var(--line);background:var(--card);cursor:pointer;transition:all .12s}
.filter-chip:hover{border-color:var(--moss)}.filter-chip.on{background:var(--moss);color:var(--paper);border-color:var(--moss)}
.filters{display:flex;gap:.5rem;flex-wrap:wrap;margin:1rem 0}
.result-count{font-size:.85rem;color:var(--ink-soft);margin:.6rem 0}
.nav-genus{display:flex;justify-content:space-between;padding:1.2rem 0;border-top:1px solid var(--line);margin-top:1rem;font-size:.88rem}
.nav-genus a{display:flex;align-items:center;gap:.4rem;color:var(--moss)}
.site-footer{border-top:1px solid var(--line);background:var(--moss);color:var(--paper-warm);padding:2.5rem 0;margin-top:3rem}
.site-footer a{color:var(--paper)}.site-footer a:hover{color:var(--gold)}
.site-footer .wrap{display:flex;justify-content:space-between;gap:2rem;flex-wrap:wrap}
.site-footer .fcol h4{font-family:var(--serif);font-size:1rem;margin-bottom:.6rem}
.site-footer .fcol a{display:block;font-size:.88rem;padding:.15rem 0}
.site-footer .fine{font-size:.78rem;opacity:.8;margin-top:1rem}
.site-footer .copyright{margin-top:.5rem}
@media(max-width:760px){.site-header .wrap{flex-direction:column;align-items:flex-start;gap:.8rem}.pheno .m{font-size:.6rem}}
.fade-up{animation:fadeUp .5s ease both}@keyframes fadeUp{0%{opacity:0;transform:translateY(12px)}to{opacity:1;transform:none}}